Herbs, subshrubs or shrubs without latex; leaves usually alternate; heads various; involucres often hemispheric, bracts in 2–7 rows, imbricate and almost always with scarious margins and apex; receptacles flat, hemispherical or conical, glabrous or hairy, paleate or ep- aleate; ray corollas white and/or yellow (rarely blue-vio- let, pinkish or reddish), limbs with three lobes; outer disc florets (if present in disciform capitula) yellow, in one to several rows, corolla with 0–5 lobes; central disc corolla yellow or rarely whitish or reddish, actinomorphic with 3–6 apical lobes; anther apex usually ovate or triangular, generally rounded, thecae usually ecalcarate and ecaudate (rarely shortly tailed); pollen echinate (rarely rugose or smooth) and spherical; style slender, apex truncate with parallel stigmatic surfaces, style hairs rather short and api- cally rounded; achenes various; pappus never capillary.

Diagnostic Features

Leaves generally variously dis- sected, rarely entire with characteristic aromatic smell; involucral bracts with hyaline margin; no tails on anthers; tubular florets hermaphrodite or functionally male, tu- bular or funnel-shaped; upper part of the filament with cells with thickened walls, forming a split cylindrical or balusterform anther collar (filament collar); and achenes without a carbonized layer, often with glandular hairs, mucilage cells and resin sacs/ducts.

Geographic distribution

Mainly in southern Africa, central Asia, and the Mediterranean region.