Shrubs or herbs without latex; leaves alter- nate; heads disciform, discoid or radiate, small to me- dium with up to 45 florets (except for Anisopappus which can have 100’s of florets); involucre cup-shaped to cy- lindrical, often much reduced, bracts imbricate without hyaline margins, in few rows; receptacle naked or paleate; corollas white or yellow; ray corolla when present, strap- shaped with three teeth; outer florets often filiform, tu- bular; disc corollas with five lobes of equal size; anther apical appendage acute, thecae generally without spurs (ecalcarate) but often with distinct tails (caudate); pol- len spinulose, caveate with one baculate layer, or with irregularly interlaced inner layer; styles slender or thick, apex slightly rounded with short obtuse sweeping hairs above where the style divides, at maturity style branches recurving; achenes without large elongated crystals; pap- pus of scales, awns or missing.

Diagnostic Features

Heads sometimes in pseudocepha- lia, some achenes have phytomelanin; and anthers with tails.

Geographic distribution

Mainly Africa and Australia