Trees, shrubs, subshrubs, or perennial herbs without latex; leaves alternate sometimes rosulate; heads solitary (or 2–3) or several, homogamous or heteroga- mous, radiate or discoid (occasionally disciform), small to large (ca. 5–300 florets); involucre oblong to cam- panulate or turbinate, bracts imbricate in 3–10 series, margins not hyaline; receptacle naked, alveolate; corollas white to orange (seldom pink, lilac, or purple); marginal florets, when present, bisexual or female, usually with staminodes, bilabiate or sub-bilabiate (rarely sub-zygo- morphic); central florets bisexual; disc corollas tubular, deeply lobed, the lobes equal in length; anther apical appendages apiculate, thecae with tails (caudate); pol- len spheroidal to prolate or elliptic, exine almost psilate; styles slender, usually shallowly divided, glabrous, apex rounded to subrounded, usually with an extension of the stigmatic surface; achenes obovate, villose; pappus of uni- seriate or 2- or 3-seriate, scabrid or occasionally plumose (at the apex) capillary bristles or less commonly some of them relatively flat.

Diagnostic Features

Marginal corollas, when present, bilabiate; disc corollas tubular, deeply lobed; anther apical appendages apiculate; and styles glabrous.

Geographic distribution

America, from southern United States to Argentina and the Caribbean (especially Cuba).