Gymnarrheneae

Amphicarpic annuals with no reports of latex; leaves alternate forming rosettes; sub- terranean heads homogamous, female, cleistogamous; flo- rets enclosed in small involucral bracts, corolla vestigial; pappus absent, vestigial, or of short, somewhat scale-like bristles; aerial heads congested in the center of the leaf rosette, heterogamous, disciform, functionally staminate florets in small groups, loosely connected on very short pedicels, interspersed among the small pistillate florets; corollas small, whitish; anther apical appendage absent, thecae without spurs (ecalcarate) and with tails (caudate); pollen echinate and non-lophate, the spines unevenly- distributed; female florets solitary, each enclosed in a prominent, stiff, white and green bract; corolla filiform; style arms long with rounded apex, glabrous, at maturity style branches not recurving; achenes of three types, those of pistillate florets numerous, tiny, ovoid, ciliate, villous, with long twin hairs; pappus of three types, that of female florets of long-lanceolate, ciliate, acuminate scales.

Diagnostic Features

Ephemeral, dwarf desert annual, amphicarpic; both chasmogamous aerial heads and cleis- togamous subterranean ones and two different types of achenes; corollas 3–4-lobed; 3–4 anthers; and aerial heads clustered together and with functional male and female florets.

Geographic distribution

Deserts in northern Africa and the Middle East.