Hecastocleideae

Small to medium shrub without latex; leaves alternate; heads discoid, small (1 floret); involucre cylindric to fusiform, bracts imbricate without hyaline margins; receptacle naked; corollas white to reddish to greenish white; disc corollas with five lobes of equal size, deeply divided; anther apical appendages lanceolate not elongate, thecae bases with spurs (calcar- ate) and slightly fimbriate tails (caudate); pollen psilate and oblong; styles slender and glabrous, branches short (0.1–0.5 mm), apices rounded, not recurving at maturity; achene terete, obscurely 4–5-nerved, glabrescent; pappus of six unequal, lanceolate or multi-toothed scales some- times fused to form lacerate crowns.

Diagnostic Features

Stiff holly-like leaves; tricolpate pollen; heads that are single-flowered and re-aggregated on a secondary receptacle, each group of 1–5 heads being subtended by a relatively large spiny whitish or greenish bract.

Geographic distribution

Restricted to high elevation areas (1500– 2000 m) in southern Nevada and adjacent California including Death Valley.