Small trees, shrubs, or subshrubs without latex; leaves alternate; heads homogamous or heterogamous, radiate or discoid, small to relatively large (5 to ca. 75 florets); involucre cylindrical to campanulate or turbinate, bracts imbricate to sub-imbricate, inner bracts longer than outer, usually in 3–10 series, green with darker or lighter margins but not hyaline, sometimes scabrous (Leucomeris); receptacle naked, flat or alveolate; florets dimorphic or isomorphic, bisexual, corollas whitish, pink, or purple, rarely yellow; marginal florets, when present, bisexual or female, sometimes with staminodes (Ianthopappus), corol- las bilabiate with lobes coiled; central florets one (Hyalis) to ca. sixty, bisexual, corollas tubular, deeply 5-lobed, the lobes equal in length and coiled; anther apical appendage apiculate, acute to acute-apiculate, thecae sagittate (spurs; calcarate) with long laciniate or papillose tails (caudate); pollen sub-prolate to prolate, elliptic, exine psilate (to nearly psilate); styles thick to slender, shallowly divided (1.5 mm), glabrous, apex rounded or subrounded, usually with an extension of the stigmatic surface (Ianthopappus), style branches never recurving; achenes fusiform, cylin- drical, ribbed, villose; pappus of 2- or 3-seriate, scabrid to smooth (occasionally plumose at the apex) capillary bristles, outer ones shorter.
Marginal corollas usually present, bilabiate; central corollas tubular, deeply lobed; anther apical appendages apiculate; styles glabrous; and pappus of 2- or 3-seriate, scabrid or occasionally plumose at the apex capillary bristles with a reduced number of outer shorter bristles.
South America and Asia.