Herbs, subshrubs, shrubs or vines without latex; leaves alternate sometimes rosulate; heads discoid or ra- diate, small to medium size; involucre cylindric to cam- panulate, bracts imbricate; receptacle naked or paleate, alveolate; corollas white, yellow, red, blue, orange, pur- ple, violet; ray corolla bilabiate with a 3-toothed outer lip and 1–2-toothed inner lip, outer lip short or long; disc corolla bilabiate (rarely with five lobes of equal size); an- ther apical appendage acute, thecae with spurs (calcarate) and tails (caudate); pollen psilate, spheroidal and subo- blate; styles thick, apex truncate, rarely rounded, arms with an apical crown of sweeping hairs; achenes elliptic to turbinate, glabrous or pubescent; pappus of capillary, paleaceous, scabrose, or plumose bristles, one to more than one series, or absent.

Diagnostic Features

All corollas bilabiate; style apex truncate, the only pubescence is on the arms, which have a unique apical crown of sweeping hairs; pollen exine distinctly bilayered, with the ectosexine and the endo- sexine clearly columellate.

Geographic distribution

Mostly southern South America.