Perennial herbs, subshrubs or shrubs, without latex; leaves alternate, often rosulate, clustered at the median part of the stem or on secondary short shoots; heads discoid, one to ca. 13(–16) florets; involucre cylin- drical to obconic, commonly multiseriate (5–15 rows) to rarely uniseriate of few involucral bracts, usually imbri- cate with hyaline margins sometimes ciliolate; receptacle epaleaceous usually glabrous, rarely pilose, alveolate or not; florets usually bisexual and fertile (rarely functionally female/male or cleistogamous), corollas white or pink, 5-lobed but zygomorphic with one split deeper than the others, sometimes pseudo-bilabiate or pseudo-ligulate; anther apical appendages truncate or rounded to apicu- late, more seldom emarginated, thecae with spurs (calcar- ate) and with tails long and smooth to pilose (caudate); pollen spheroidal or subprolate, exine microechinate, scabrous, or echinate; styles slender, branches bilobed to shortly branched, variously truncate, rounded or acute at the apex, dorsally pilose to shortly pilose, sometimes swelling slightly on upper part of style with branches recurved; achenes oblong to obovate, (9- or) 10-veined, glabrous or pilose; pappus bristles 1–3-seriate (sometimes absent in functionally male or chasmogamous florets), margins scabrid or plumose.

Diagnostic Features

Small discoid heads of white, pink, rose or purple florets; corollas irregularly deeply 5-lobed and therefore zygomorphic by having one sinus deeper than the others, appearing pseudo-bilabiate or pseudo- ligulate; and styles thick, branches bilobed to shortly branched, dorsally pilose to shortly pilose.

Geographic distribution

Southeast Asia